International Journal of Myriapodology 4: 51–77, doi: 10.3897/ijm.4.1103

New records of Pauropoda (Myriapoda) from north-western Thailand

| |

(C) 2011 Ulf Scheller. This is an open access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.

For reference, use of the paginated PDF or printed version of this article is recommended.

Abstract

Fourteen species of Pauropoda are reported from
altitudes above 1000 m in north-western Thailand, among them one
species belonging to a new genus in Pauropodidae, *Angkapauropus* **gen. n.**, and one species in *Borneopauropus*, a genus in Brachypauropodidae, a family not previously reported from Thailand. The species new to science are: *Decapauropus prolixus* **sp. n.**, *Decapauropus capillaris* **sp. n.**, *Decapauropus brevilingulus* **sp. n.**, *Decapauropus premnos* **sp. n.**, *Decapauropus anatanosus* **sp. n.**, *Decapauropus undulatulus* **sp. n.**, and *Angkapauropus leptotrichos* gen. n., **sp. n.** (all in Pauropodidae); *Borneopauropus platylopas* **sp. n.** (Brachypauropodidae); *Samarangopus choanephorus* **sp. n.** (Eurypauropodidae), and *Sphaeropauropus angulatus* **sp. n.** (Sphaeropauropodidae).

Keywords

Myriapoda, Pauropoda, new genus, new species, taxonomy, Thailand, biogeography

Introduction

Genus*Decapauropus
* Remy, 1931
*Decapauropus*
*grandicollis*
Scheller, 1995
*Decapauropus*
*prolixus*
sp. n.
*Decapauropus*
*capillaris*
sp. n.
*Decapauropus*
*brevilingulus*
sp. n.
*Decapauropus*
*premnos*
sp. n.
*Decapauropus*
*anatonus*
sp. n.
*Decapauropus*
*undulatulus*
sp. n.
*Angkapauropus*
*leptotrichos*
sp. n.
Family Brachypauropodidae Silvestri, 1902

Genus*Borneopauropus*
Scheller, 2008
*Borneopauropus*
*platylopas*
sp. n.
Family Eurypauropodidae Ryder, 1879

Genus*Samarangopus*
Verhoeff, 1934
*Samarangopus*
*poculifer*
Scheller, 1995
*Samarangopus*
*umbonifer*
Scheller, 1995
*Samarangopus*
*choanephorus*
sp. n.
Family Sphaeropauropodidae Verhoeff, 1934 Genus Sphaeropauropus Silvestri, 1930
*Sphaeropauropus*
*convolvolutus*
Scheller, 1995
*Sphaeropauropus*
*angulatus*
sp. n.
Remarks

Pauropoda were first recorded from Thailand by *Allopauropus* and one in *Decapauropus*, from the Island Koh Chang in the Gulf of Thailand. The present author then reported 11 species, seven in *Decapauropus*, three in *Samarangopus* and one in *Sphaeropauropus*, in a collection from Doi Inthanon in the northwest (

All specimens were collected from Berlese funnel extractions and were preserved in ethanol and studied in monopropylene glycol. In the descriptions below individuals have been classified as adults, subadults and juveniles according to the number of pairs of legs. The sex of adults and subadults was recorded.

The type material has been lodged in the collections of the Zoological Museum, University of Lund, Sweden.

Systematics Family Pauropodidae Verhoeff, 1934Genus

Material.

Thailand, Chiang Mai province, Doi Inthanon, Mae Chaem road, secondary dry forest, litter, alt. 1150 m, 1 juv. 5, 1991.vi.30, loc. CM-080. – 1 specimen.

General distribution.

Known previously from Doi Inthanon only (

urn:lsid:zoobank.org:act:BBC206B8-80B2-4D40-96C5-F1BA62CF0E9F

http://species-id.net/wiki/Decapauropus_prolixus

Figs 1–9
Material.

Holotype ad. 9(♂), Thailand, Chiang Mai province, Doi Inthanon, Mae Chaem road, secondary dry forest, litter, alt. 1150 m, 1991.vii.4, loc. CM-096. – 1 specimen.

Etymology.

From the Latin *prolictus*, stretched out long (referring to the posterolateral appendages of the anal plate).

Diagnosis.

*Decapauropus prolixus* may be close to *Decapauropus bispinus* *Decapauropus prolixus*, large in *Decapauropus bispinus*, the antennal globulus *g*, pyriform, not spherical, the shape of the pygidial *st*,
long, thin, cylindrical, not short, globular, and the lateral
margins of the anal plate, parallel, not distinctly convex.

Description.

*Length.* 0.48 mm. *Head* (Fig. 1). Setae on the tergal side subcylindrical striate. Relative lengths of setae, 1^{st} row: *a*_{1}=10, *a*_{2}=12; 2^{nd} row: *a*_{1}=14, *a*_{2}=15, *a*_{3}=17; 3^{rd} row: *a*_{1}=12, *a*_{2}=15; 4^{th} row: *a*_{1}=14, *a*_{2}*=*15, *a*_{3}=18, *a*_{4}=11; lateral group setae, *l*_{1}=18, *l*_{2}≈17, *l*_{3}=?. Ratio *a*_{1}*/a*1*-a*1
in 1^{st} row 0.9, 2^{nd} row 0.5, 3^{rd} row 0.8, 4^{th} row 1.5. Temporal
organs small, in tergal view ovoid, as long as their shortest distance
apart. Head cuticle glabrous.

*Antennae* (Fig. 2). Segment 4 with 4 cylindrical striate setae; their relative lengths: *p*=10, *p’*=8, *p’’*=2, *r*=5. Tergal seta *p* 0.9 of the length of tergal branch *t*. The latter fusiform, 2.7 times as long as its greatest diameter and 1.1 times as long as the length of sternal branch*s*, that branch 1.7 times as long as its greatest diameter; anterodistal corner distinctly truncate. Seta *q* cylindrical striate, 0.7 of the length of *s*. Relative lengths of flagella (basal segments included) and basal segments: *F*_{1}=100, *bs*_{1}=7; *F*_{2}=58, *bs*_{2}=5; *F*_{3}=91, *bs*_{3}=5. *F*_{1} 3.4 times as long as *t*, *F*_{2} and *F*_{3} 2.1 and 3.3 times as long as *s* respectively. Distal calyces of *F*_{1} and *F*_{3} conical, those of *F*_{2} hemispherical, distal part of flagella axes inconsiderably widened below calyces. Globulus *g* proportionally large, pyriform, 1.5 times as long as wide, ≈10 bracts, capsule subspherical; width of *g* as long as the greatest diameter of *t*. Antennae glabrous.

*Trunk* (Figs 3, 4). Setae of collum segment (Fig. 3)
simple cylindrical striate, sublateral setae 1.3 times as long as
submedian setae; sternite process triangular, blunt anteriorly;
appendages tapering distally, caps low. Process and appendages
glabrous. Setae on tergites as setae on the head, 4+4 setae on tergite I
and ?4+2 on VI , interposed tergites not studied. Submedian posterior
setae on VI (Fig. 4) 0.5 of interdistance and 0.6 of the length of pygidial setae *a*_{1}.

*Genital papillae* (Fig. 5).
Proximal 2/3 subcylindrical, distal part tapering into an
subcylindrical top, papillae 2.1 times as long as the greatest
diameter, seta thin, 0.3 of the length of papilla.

*Bothriotricha* (Fig. 6). Relative lengths: *T*_{1}=*T*_{3}=100, *T*_{2}=108, *T*_{4}=138, *T*_{5}=196; axes simple straight, very thin and with faint pubescence only distally, *T*_{3} only (Fig. 6) stronger, fusiformly thickened in distal ¾ and with distinct oblique pubescence.

*Legs* (Figs 7, 8). Setae on coxa and trochanter (Fig. 7)
of leg 9 furcate, branches cylindrical striate, secondary branch
shorter than primary one. These setae more anteriorly with rudimentary
secondary branches except in coxal setae of leg 2. Tarsus of leg 9 (Fig. 8)
tapering, 3.5 times as long as its greatest diameter; proximal seta
very thin tapering pointed, distal seta subcylindrical blunt striate,
proximal one 0.3 of the length of tarsus and as long as the length of
distal seta. Cuticle of tarsus glabrous.

*Pygidium* (Fig. 9). *Tergum*. Posterior margin between *st* straight. Relative lengths of setae: *a*_{1}=10, *a*_{2}=9, *a*_{3}=19, *st*=6; setae directed posteriorly, *a*_{1} and *a*_{3} almost straight, *a*_{2} and *st* curved inwards, *a*_{1} striate distally. Distance *a*_{1}*-a*1 0.9 of the length of *a*_{1}, distance *a*_{1}*-a*2 twice longer than distance *a*_{2}*-a*3; distance *st-st* 1.3 times as long as *st* and as long as distance *a*_{1}*-a*1.

*Sternum*. Posterior margin with low and broad bulge below base of anal plate. Relative lengths of setae (pygidial *a*_{1}=10): *b*_{1}=31, *b*_{2}=10, setae thin tapering, *b*_{1} with short pubescence distally, 1.4 times as long as interdistance, *b*_{2} 0.7 of distance *b*_{1}-*b*_{2}.

Anal plate directed posteriorly, lateral margins parallel, posterodistal corners lengthened into two long tapering and distally faintly striate appendages, these ≈1.6 times as long as the length of plate; posterior margin between appendages U-shaped and with two short tapering glabrous appendages protruding backwards from sternal margin, length of short appendages ≈¼ of the length of posterolateral appendages; plate glabrous.

Figures 1–9.*Decapauropus prolixus* sp. n., holotype, ad. 9(♂) **1** head, median and right part **2** left antenna, sternal view **3** collum segment, median and left part, sternal view **4** tergite VI, posterior part **5** genital papillae and seta on coxa of leg 2, anterior view; **6** *T*_{3}**7** seta on trochanter of leg 9 **8** tarsus of leg 9 **9** pygidium, posteromedian and left part, sternal view. Scale: a: Figs 4–8; b: 1–3; c: 9.

urn:lsid:zoobank.org:act:A56C4313-659B-42AB-8FFF-540764DB2D29

http://species-id.net/wiki/Decapauropus_capillaris

Figs 10–19
Material.

Holotype ad. 9(♂), Thailand, Chiang Mai province, Doi Inthanon, Mae Chaem road, secondary dry forest, litter, alt. 1150 m, 1991.vi.30, loc. CM-080. – Paratypes: 3 ad. 9 (2♀, 1♂), same data as holotype. – 4 specimens.

Etymology.

From the Latin *capillaris*, slender as a hair (referring to the thin setae).

Diagnosis.

*Decapauropus capillaris* shows striking resemblances to *Decapauropus virgininsularis* Scheller (*Decapauropus capillaris*,
not 4.9–5.3, the setae of the collum segment, both of the same
shape, not sublateral seta much longer and thicker than submedian one,
the tarsus of the last pair of legs, only somewhat tapering, not
strongly tapering and very slender distally, the size of the *st*, rudimentary, not well developed, and the shape of the lateral margins of the anal plate, convex, not concave.

Description.

*Length.* (0.50-)0.52(-0.65) mm. *Head* (Fig. 10). Setae on the tergal side thin striate. Relative lengths of setae (holotype only), 1^{st} row: *a*_{1}=*a*_{2}=10; 2^{nd} row: *a*_{1}=11, *a*_{2}=13, *a*_{3}=15; 3^{rd} row: *a*_{1}=?, *a*_{2}=11; 4^{th} row: *a*_{1}=?, *a*_{2}*=*14, *a*_{3}=16, *a*_{4}=?; lateral group setae, *l*_{1}=19, *l*_{2}=17, *l*_{3}=10. Ratio *a*_{1}*/a*1*-a*1
in 1^{st} row 1.0, 2^{nd} row 0.8, 3^{rd} and 4^{th} row?. Temporal organs in
tergal view broadest in posterior part, length 0.9 of their shortest
distance apart. Pore not ascertained. Head cuticle glabrous.

*Antennae* (Fig. 11). Segment 4 with five setae, all thin cylindrical striate; their relative lengths: *p*=10, *p’*=44, *p’’*=11, *p’’’*=5, *r*=39. Tergal seta *p* 1.2 times as long as the length of tergal branch *t*. The latter fusiform, (3.4-)3.5 times as long as its greatest diameter and 1.3 times as long as the length of sternal branch*s*, that branch 2.1 times as long as its greatest diameter; anterodistal corner distinctly truncate. Seta *q* thin cylindrical striate, 1.3 times as long as the length of *s*. Relative lengths of flagella (basal segments included) and basal segments: *F*_{1}=100, *bs*_{1}=5; *F*_{2}=?, *bs*_{2}=4; *F*_{3}=76, *bs*_{3}=5. *F*_{1} 3.7 times as long as *t*, *F*_{3} 3.0 times as long as *s*. Distal calyces small rounded, distal part of flagella axes not widened below calyces. Globulus *g* subspherical, 1.2 times as long as wide, 8 bracts, capsule with flattened bottom; width of *g* 0.8 of the greatest diameter of *t*. Antennae glabrous.

*Trunk* (Figs 12, 13). Setae of collum segment (Fig. 12)
furcate, main branch cylindrical blunt striate, secondary branch
rudimentary, sublateral setae 1.7 times as long as submedian setae;
sternite process triangular, pointed anteriorly; appendages tapering
distally. Process and appendages glabrous.

Setae on tergites as setae on the head, 4+4 setae on tergite I, 6+6 on II-V, 4+2 on VI. Submedian posterior setae on VI (Fig. 13) long tapering pointed, 1.4 times as long as interdistance.

*Genital papillae* (Fig. 14). Conical, 2.0 times as long as the greatest diameter, seta thin, 0.4 of the length of papilla.

*Bothriotricha* (Fig. 15). Relative lengths (holotype only): *T*_{1}=100, *T*_{2}=152, *T*_{3}=105, *T*_{4}=123, *T*_{5}=131; axes simple straight, very thin and with short pubescence, erect distally; *T*_{3} only (Fig. 15) stronger, fusiformly thickened in proximal ¾ and with distinct oblique pubescence.

*Legs* (Figs 16, 17). Setae on coxa and trochanter (Fig. 16)
of leg 9 furcate, branches cylindrical striate, secondary branch
shorter than primary one. These setae more anteriorly with rudimentary
secondary branches except in coxal setae of leg 2 (paratype male).
Tarsus of leg 9 (Fig. 17)
tapering, 3.5(-3.6) times as long as its greatest diameter; setae
thin, proximal one tapering pointed, distal one subcylindrical blunt,
faintly striate, proximal seta (0.3-)0.4 of the length of tarsus and
(3.4-)3.5 times as long as distal seta. Cuticle of tarsus glabrous.

*Pygidium* (Figs 18, 19). *Tergum*. Posterior margin rounded but with small posterior lobe between *st*. Setae *a*_{1} broken or rudimentary, relative lengths of setae: *a*_{2}=10, *a*_{3}=18, *st* rudimentary; setae directed posteriorly, *a*_{2} and *a*_{3} long thin tapering glabrous, somewhat curved inward. Distance *a*_{1}*-a*2 three times longer than distance *a*_{2}*-a*3; distance *st-st* ≈20 times as long as *st* and 0.4 of distance *a*_{1}*-a*1.

*Sternum*. Posterior margin with distinct indentation and broadly triangular lobe below anal plate. Relative lengths of setae (pygidial *a*_{2}=10): *b*_{1}=7, *b*_{2}=3, setae thin tapering, *b*_{1} indistinctly striate in distal half, as long as interdistance, *b*_{2} 0.7(-0.8) of distance *b*_{1}-*b*_{2}.

Anal plate somewhat turned up, narrowest anteriorly, linguiform, lateral margins a little convex, on underside of rounded distal part two very short cylindrical blunt appendages protruding backwards-downwards; plate glabrous.

Figures 10–19.

*Decapauropus capillaris* sp. n., holotype, ad. 9(♂) **10** head, median and right part **11** right antenna, sternal view **12** collum segment, median and left part, sternal view **13** tergite VI, posteromedian part and right posterior corner **14** genital papillae, anterior view **15** *T*_{3}**16** seta on coxa of leg 9 **17** tarsus of leg 9 **18** pygidium, posteromedian and left part, sternal view **19** anal plate, lateral view. Scale: a: Figs 13–17; b: 10, 12, 18, 19; c: 11.

urn:lsid:zoobank.org:act:FEEA6B7A-30E5-43CB-9027-3ECA9438AA63

http://species-id.net/wiki/Decapauropus_brevilingulus

Figs 20–27
Material.

Holotype ad. 9(♀), Thailand, Chiang Mai province, Doi Inthanon, below the top, alt. 2400 m, 1991.vii.8, loc. CM-243.

Etymology.

From the Latin *brevis*, short, and *lingua*, tongue (referring to the shape of the anal plate).

Diagnosis.

*Decapauropus brevilingulus*
sp. n. is well defined by its unique short linguiform anal plate in
combination with a distinct semicircular lobe between the pygidial setae
*st* and longish antennal globulus with ovoid capsule. At present its relationships can not be traced.

Description.

*Length.* 0.55 mm. *Head* (Fig. 20). Setae on the tergal side thin annulate. Relative lengths of setae, 1^{st} row: *a*_{1}=10, *a*_{2}=?; 2^{nd} row: *a*_{1}=10, *a*_{2}=11, *a*_{3}=14; 3^{rd} row: *a*_{1}=?, *a*_{2}=10; 4^{th} row: *a*_{1}=9, *a*_{2}*=a*4=11, *a*_{3}=13; lateral group setae not studied. Ratio *a*_{1}*/a*1*-a*1
in 1^{st} row 1.4, 2^{nd} row 0.5, 3^{rd} row?, 4^{th} row 1.3. Temporal organs
small, ovoid in tergal view, length 0.6 of their shortest distance
apart; small pore near posterior margin. Head cuticle glabrous.

*Antennae* (Fig. 21). Segment 4 with at least four setae, all thin cylindrical annulate blunt; their relative lengths: *p*=10, *p’*=7, *p’’*=6, *r*=5. Tergal seta *p* 0.8 of the length of tergal branch *t*. The latter proportionally short, somewhat fusiform, twice longer than thick and 0.8 of the length of sternal branch*s*, that branch 1.9 times as long as its greatest diameter; anterodistal corner distinctly truncate. Seta *q* thin cylindrical annulate, ≈0.5 of the length of *s*. Relative lengths of flagella (basal segments included) and basal segments: *F*_{1}=100, *bs*_{1}=5; *F*_{2}=38, *bs*_{2}=6; *F*_{3}=85, *bs*_{3}=8. *F*_{1} 5.5 times as long as *t*, *F*_{2} and *F*_{3} 1.6 and 3.6 times as long as *s* respectively. Distal calyces somewhat flattened, distal part of flagella axes widened fusiformly below calyces. Globulus *g* longish, 1.5 times as long as greatest diameter, bracts and capsule proportionally long, 7 bracts; *g* as wide as the greatest diameter of *t*. Antennae glabrous.

*Trunk*. Setae of collum segment (Fig. 22)
simple cylindrical blunt striate, sublateral and submedian setae of
the same length; sternite process triangular, pointed anteriorly;
appendages subspherical with flat caps. Process and appendages glabrous.

Most setae on tergites not available for study, those studied similar to those on head, 4+4 setae on tergite I, 4+2 on VI.

*Bothriotricha*. Most bothriotricha broken; *T*_{3} (Fig. 23) with proximal 2/3 somewhat thickened, pubescence distinct, oblique on thickened part, erect distally; *T*_{5} thin with very faint oblique pubescence.

*Legs* (Figs 24, 25). Setae on coxa (Fig. 24) and trochanter of all legs simple cylindrical striate. Tarsus of leg 9 (Fig. 25)
tapering, 4.8 times as long as its greatest diameter; setae thin,
proximal one curved tapering pointed glabrous, distal one
subcylindrical blunt striate, proximal seta 0.2 of the length of tarsus
and 1.3 times as long as distal seta. Cuticle of tarsus glabrous.

*Pygidium* (Figs 26, 27). *Tergum*. Posterior margin with large semicircular lobe between *st*. Setae *a*_{2} and *a*_{3} distinctly shorter than *a*_{1}; relative lengths of setae: *a*_{1}=10, *a*_{2}=6, *a*_{3}=7, *st*=3; *a-*setae directed posteriorly, proportionately long, tapering glabrous; *a*_{1} straight, *a*_{2} and *a*_{3} curved inward. Distance *a*_{1}*-a*1 as long as *a*_{1}, distance *a*_{1}*-a*2 3.3 times as long as distance *a*_{2}*-a*3; distance *st-st* 1.8 times as long as *st* and 0.8 of distance *a*_{1}*-a*1.

*Sternum*. Posterior margin with shallow indentation between *b*_{1}. Relative lengths of setae (pygidial *a*_{1}=10): *b*_{1}=16, *b*_{2}=7, setae thin tapering, *b*_{1} indistinctly striate in distal half, 1.2 times as long as interdistance, *b*_{2} as long as distance *b*_{1}-*b*_{2}.

Anal plate horizontal, narrowest anteriorly, somewhat broader than long, shortly linguiform, distal margin somewhat thickened at posterior margin; no appendages.

Figures 20–27.

*Decapauropus brevilingulus* sp. n., holotype, ad. 9(♂) **20** head, median and right part **21** left antenna, sternal view **22** collum segment, median and left part, sternal view **23** *T*_{3}**24** seta on coxa of leg 9 **25** tarsus of leg 9 **26** pygidium, posteromedian and left part, sternal view **27** anal plate, lateral view. Scale: a: Fig. 23; b: Fig. 20; c: 21, 22, 24, 25; d: 26, 27.

urn:lsid:zoobank.org:act:DEE020F4-FABC-4962-B5FF-F52E9BEDF175

http://species-id.net/wiki/Decapauropus_premnos

Figs 28–35
Material.

Holotype ad. 9(♀), Thailand, Chiang Mai province, Doi Inthanon, below the top, primary rain forest, soil, alt. 2400 m, 1991.vii.4, loc. CM-195. – Paratype subad. 8(♀), below the top, primary rain forest, soil, alt. 2400 m, 1991.vii.8, loc. CM-243. – Other material, 1 juv. 5, ibidem, Mae Chaem road, secondary dry forest, litter, alt. 1150 m, 1991.vi.30, CM-064. – 3 specimens.

Etymology.

From the Greek *premnos*, stump of a tree (referring to the thick stalk of the antennal globulus *g*).

Diagnosis.

*Decapauropus premnos* seems to be close to *Decapauropus mortensenii*
(Hansen) which was described (1902) from the island Koh Chang in the
Gulf of Thailand and later has been reported from Egypt, Seychelles,
Sri Lanka, Java and Sumatra, and New Caledonia (with uncertainty also
from Réunion, Mauritius and Australia). The new species can be
distinguished from *Decapauropus mortensenii* by its thick stalk of the antennal globulus *g*, narrow in *Decapauropus mortensenii*, the distal part of the bothriotrix *T*_{3},
with marked swelling, not evenly thickened, by the shape of the
tarsi, subcylindrical and only somewhat tapering, not slender and
strongly tapering, by the proportion *b*_{2}/*b*_{1}-*b*_{2}, as long as, not *b*_{2}>*b*_{1}-*b*_{2},
and by the shape of the anal plate, longish and with evenly convex
lateral margins, not shorter/broader and with shallow lateral
indentation.

Description.

*Length.* 0.62 mm. *Head* (Fig. 28). Setae on the tergal side thin cylindrical annulate. Relative lengths of setae, 1^{st} row: *a*_{1}=*a*_{2}=10; 2^{nd} row: *a*_{1}=?, *a*_{2}=12, *a*_{3}=14; 3^{rd} row: *a*_{1}=11, *a*_{2}=10; 4^{th} row: *a*_{1}=8, *a*_{2}*=*10, *a*_{3}=?, *a*_{4}=11; lateral group setae, *l*_{1}=25, *l*_{2}=26, *l*_{3}=?. Ratio *a*_{1}*/a*1*-a*1
in 1^{st} row 1.0, 2^{nd} row ?, 3^{rd} row 0.6, 4^{th} row 0.8. Temporal organs
large, in tergal view ovoid, length 0.8 of shortest interdistance.
Head cuticle glabrous.

*Antennae* (Fig. 29). Segment 4 with at least three cylindrical annulate setae; their relative lengths: *p*=10, *p’*=6, *r*=5. Tergal seta *p* 1.2 times as long as the length of tergal branch *t*.
The latter fusiform with distal end cut obliquely, 2.1 times as long
as its greatest diameter and 0.9 of the length of sternal branch*s*, that branch twice longer than its greatest diameter; anterodistal corner distinctly truncate. Seta *q* cylindrical striate, 0.6 of the length of *s*. Relative lengths of flagella (basal segments included) and basal segments: *F*_{1}=100, *bs*_{1}=6; *F*_{2}=27, *bs*_{2}=4; *F*_{3}=82, *bs*_{3}=5. *F*_{1} 5.0 times as long as *t*, *F*_{2} and *F*_{3} 1.2 and 3.6 times as long as *s* respectively. Distal calyces somewhat flattened, distal part of flagella axes inconsiderably widened below calyces. Globulus *g*
proportionally large, pyriform, with thick stalk, 1.2 times as long
as wide, ≈12 thin bracts, capsule with flattened bottom; width of *g* 1.1 times as long as the greatest diameter of *t*. Antennae glabrous.

*Trunk* (Figs 30, 31). Setae of collum segment (Fig. 30)
simple cylindrical striate, sublateral setae 1.4 times as long as
submedian setae; sternite process triangular, blunt anteriorly;
appendages barrel-shaped, caps low. Process and appendages glabrous.
Setae on tergites as setae on the head, 4+4 setae on tergite I and 4+2
on VI, interposed tergites not studied. Submedian posterior setae on VI
(Fig. 31) 0.3 of interdistance and 0.8 of the length of pygidial setae *a*_{1}.

*Bothriotricha* (Fig. 32). Relative lengths: *T*_{1}=100, *T*_{2}=103, *T*_{3}=76, *T*_{4}=?, *T*_{5}=107; axes simple straight, very thin and with faint pubescence only distally on *T*_{1}, *T*_{2} and *T*_{5}, *T*_{3} only (Fig. 32) stronger, with longish cylindrical end-swelling and distinct oblique pubescence.

*Legs* (Figs 33, 34). Setae on coxa (Fig. 33)
and trochanter of leg 9 furcate, branches thin cylindrical striate,
secondary branch somewhat thinner and shorter than primary one. These
setae more anteriorly with rudimentary secondary branches. Tarsus of leg
9 (Fig. 34)
subcylindrical, only somewhat tapering, 2.9 times as long as its
greatest diameter; setae thin, proximal one tapering pointed, distal
one subcylindrical blunt striate, proximal seta 0.4 of the length of
tarsus and 2.1 times as long the length of distal seta. Cuticle of
tarsus glabrous.

*Pygidium* (Fig. 35). *Tergum*. Posterior margin straight but with posterolateral corners rounded and two small rounded lobes just outside *st.* Relative lengths of setae: *a*_{1}=*st*=10, *a*_{2}=16, *a*_{3}=26. Setae curved inwards, *st* also converging, *a*_{1} and *st* cylindrical with small swelling distally, *a*_{2} and *a*_{3} long thin tapering, *a*_{2} pointed. Distance *a*_{1}*-a*1 0.7 of the length of *a*_{1}, distance *a*_{1}*-a*2 1.4 times as long as *a*_{2}*-a*3; distance *st-st* 1.8 times as long as *st* and 1.1 times as long as distance *a*_{1}*-a*1.

*Sternum*. Posterior margin with five rounded lobes, one on each side of insertion areas of setae *b*_{1} and one below base of anal plate. Relative lengths of setae (pygidial *a*_{1}=10): *b*_{1}=28, *b*_{2}=19, setae thin tapering pointed glabrous, *b*_{1} 1.1 times as long as interdistance, *b*_{2} 1.1 times as long as distance *b*_{1}-*b*_{2}.

Anal plate large, directed posteriorly, 1.7 times longer than broad, lateral margins convex, narrowing off posteriorly and with U-shaped posterior incision, length of the latter 0.3 of the length of plate, plate glabrous.

Figures 28–35.

*Decapauropus premnos* sp. n., holotype, ad. 9(♀) **28** head, median and right part **29** right antenna, tergal view **30** collum segment, median and left part, sternal view **31** tergite VI, posterior part **32** *T*_{3}**33** seta on coxa of leg 9; **34** tarsus of leg 9 **35** pygidium, posteromedian and left posterior corner, tergal view. Scale: a: Figs 32–34; b: 28, 29, 31; c: 30. 35.

urn:lsid:zoobank.org:act:02DDC8EF-3D76-4250-9FCC-50680FD665CE

http://species-id.net/wiki/Decapauropus_anatonus

Figs 36–45
Material.

Holotype ad. 9(♂), Thailand, Chiang Mai province, Doi Inthanon, below the top, primary rain forest, litter, alt. 2400 m, 1991.vii.4, loc. CM-210.

Etymology.

From the Latin *anatonus*, extending upward (referring to the shape of the distal part of the bracts of the antennal globulus *g*).

Diagnosis.

There are some species with resembling though not identical anal plates but *Decapauropus anatonus* sp. n. is well defined by the shape of the antennal globulus *g*
with slender stalk and distal parts of the bract turned upward, this
also combined with a distal swelling on the bothriotricha *T*_{3}. At present its relationships are not possible to trace.

Description.

*Length.* 0.43 mm. *Head* (Fig. 36). Setae on the tergal side subcylindrical striate. Relative lengths of setae, 1^{st} row: *a*_{1}=*a*_{2}=10; 2^{nd} row: *a*_{1}=10, *a*_{2}=14, *a*_{3}=20; 3^{rd} row: *a*_{1}=11, *a*_{2}=?; 4^{th} row: *a*_{1}=10, *a*_{2}*=*17, *a*_{3}=24, *a*_{4}=13; lateral group setae not studied. Ratio *a*_{1}*/a*1*-a*1
in 1^{st} row 1.4, 2^{nd} row 0.5, 3^{rd} row 1.3, 4^{th} row 1.2. Temporal
organs in tergal view ovoid, length 0.8 of their shortest
interdistance. Head cuticle glabrous.

*Antennae* (Fig. 37). Segment 4 with at least three cylindrical striate setae; their relative lengths: *p*=10, *p’*=*r*=7. Tergal seta *p* 1.5 times as long as tergal branch *t*. The latter fusiform, twice longer than its greatest diameter and 0.7 of the length of sternal branch*s*, that branch 1.8 times as long as its greatest diameter; anterodistal corner distinctly truncate. Seta *q* cylindrical striate, 0.6 of the length of *s*. Relative lengths of flagella not available for study; *F*_{1} long, distal calyces flattened, distal part of flagella axes inconsiderably widened below calyx. *F*_{1} 6.2 times as long as *t*. Globulus *g*
pyriform, proportionally long, 1.7 times as long as the greatest
diameter, stalk as long as globulus, bracts thin, turned upward
distally, ≈10 bracts, capsule with flattened bottom, width of g 0.8
of the greatest diameter of *t*. Antennae glabrous.

*Trunk* (Figs 38, 39). Setae of collum segment (Fig. 38)
simple cylindrical striate, sublateral setae 1.3 times as long as
submedian setae; sternal process blunt anteriorly, lateral margins
concave; caps low, roundly conical. Process and appendages glabrous.

Tergite I incompletely divided transversally;
setae on tergites as setae on the head, 4+4 setae on tergite I and 4+2
on VI, interposed tergites not studied. Submedian posterior setae on
VI (Fig. 39) 0.5 of interdistance and 0.8 of the length of pygidial setae *a*_{1}.

*Genital papillae* (Fig. 40). Conical, 1.9 times as long as the greatest diameter, seta thin, 0.5 of the length of papilla.

*Bothriotricha* (Fig. 41). Relative lengths: *T*_{1}=100, *T*_{2}=?, *T*_{3}=112, *T*_{4}=140, *T*_{5}=157; axes simple straight, very thin and with faint pubescence only distally, *T*_{3} (Fig. 41) strongest, with narrowly ovoid end-swelling and with distinct oblique pubescence on swelling and just below it.

*Legs* (Figs 42, 43). Setae on coxa and trochanter (Fig. 42) of legs 1–9 simple cylindrical striate. Tarsus of leg 9 (Fig. 43)
tapering, 3.6 times as long as its greatest diameter; proximal seta
very thin tapering, distal seta cylindrical blunt striate, proximal
one 0.2 of the length of tarsus and 1.2 times as long as the length of
distal seta. Cuticle of tarsus glabrous.

*Pygidium* (Fig. 44, 45). *Tergum*. Posterior margin rounded and with shallow lobe between *st*. Relative lengths of setae: *a*_{1}=10, *a*_{2}=8, *a*_{3}=17, *st*=6; setae thin, directed posteriorly, *a*-setae tapering pointed, curved inward, *st* cylindrical blunt. Distance *a*_{1}*-a*1 0.6 of the length of *a*_{1}, distance *a*_{1}*-a*2 1.8 times as long as *a*_{2}*-a*3; distance *st-st* 1.8 times as long *st* and twice longer than distance *a*_{1}*-a*1.

*Sternum*. Posterior margin rounded and with very small posterior lobe below anal plate. Relative lengths of setae (pygidial *a*_{1}=10): *b*_{1}=32, *b*_{2}=8, setae thin tapering, striate, *b*_{1} 1.3 times as long as interdistance, *b*_{2} 0.5 of distance *b*_{1}-*b*_{2}.

Anal plate directed posteriorly, as long as broad, lateral margins convex, posterior margin with shallow incision, posterodistal corners lengthened into two clavate appendages, these 0.7 of the length of plate; plate glabrous.

Figures 36–45.

*Decapauropus antanosus* sp. n., holotype, ad. 9(♂) **36** head, median and right part **37** right antenna, tergal view **38** collum segment, median and left part, sternal view **39** tergite VI, posterior part **40** genital papillae, anterior view **41** *T*_{3}**42** seta on trochanter of leg 9 **43** tarsus of leg 9 **44** pygidium, posteromedian and right part, tergal view **45** anal plate, lateral view. Scale: a: Fig. 41; b: 36, 38, 40, 42, 43; c: 37, 39, 44, 45.

urn:lsid:zoobank.org:act:05C6DFE9-0C2A-4742-8208-EF344964B483

http://species-id.net/wiki/Decapauropus_undulatulus

Figs 46–52
Material.

Holotype ad. 9(♀), Thailand, Chiang Mai province, Doi Inthanon, Mae Chaem road, secondary dry forest, litter, alt. 1150 m, 1991.vi.30, loc. CM-080.

Etymology.

From the Latin *undulatus*, wavy (referring to the distal margin of the 4^{th} antennal segment).

Diagnosis.

Many species in the genus have linguiform
anal plates with two appendages but none of them has the appendages
placed in the middle of the tergal side as in *Decapauropus undulatus*
sp. n. Moreover is this species well defined by the peculiar undulated
distal margin of the 4^{th} antennal segment and the wide collum appendages
which are directed inward/anteriorly. At present its relationships are
not possible to trace.

Description.

*Length.* 0.52 mm. *Head*. Not available for study.

*Antennae* (Fig. 46). Distal margin of segment 4 distinctly wavy, four setae; their relative lengths: *p*=10, *p’*=7, *p’’*=3, *r*=4. Setae annulate, *p* and *p’* grow thicker outward, *p’’* and *r* cylindrical. Tergal seta *p* 1.5 times as long as the length of tergal branch *t*. The latter fusiform, 1.3 times as long as its greatest diameter and 0.8 of the length of sternal branch*s*, that branch subcylindrical, 1.3 times as long as its greatest diameter; anterior truncation weak. Seta *q* cylindrical striate, inserted very near distal margin, as long as the length of *s*. Relative lengths of flagella (basal segments included) and basal segments: *F*_{1}=100, *bs*_{1}=7; *F*_{2}=44, *bs*_{2}=4; *F*_{3}=88, *bs*_{3}=7. *F*_{1} 5.5 times as long as *t*, *F*_{2} and *F*_{3} 4.6 and 5.1 times as long as *s* respectively. Distal calyces small, distal part of flagella axes inconsiderably widened below calyces. Globulus *g* proportionally large, pyriform, 1.4 times as long as wide, ≈9 bracts, capsule with flattened bottom; width of *g* 0.6 of the length of the greatest diameter of *t*. Antennae glabrous.

*Trunk* (Figs 47, 48). Setae on collum segment simple cylindrical striate, only one pair could be proved (Fig. 47);
sternite process small, indistinct; appendages large short cylindrical
directed inward and anteriorly, caps low. Process and appendages
glabrous.

Setae on tergites only partly available for study, those proved thin cylindrical. Tergite VI with 4+2 setae, submedian posterior ones (Fig. 48) 0.9 of interdistance.

*Bothriotricha* (Fig. 49). Relative lengths: *T*_{1}=100, *T*_{2}=98, *T*_{3}=79, *T*_{4}=104, *T*_{5}=?; axes simple straight thin in *T*_{1}, *T*_{2}, *T*_{4} and *T*_{5}, pubescence there faint on proximal halves, on distal halves of *T*_{1}, *T*_{2} and *T*_{4} distinct, partly erect, probably branched most distally; axes of *T*_{3} (Fig. 49) somewhat thickened in proximal half, pubescence there oblique, on distal half as on other bothriotricha.

*Legs* (Figs 50, 51). Setae on coxa and trochanter (Fig. 50)
of leg 9 furcate, branches cylindrical blunt striate, secondary
branch shorter and thinner than primary branch. These setae more
anteriorly with rudimentary secondary branches. Tarsus of leg 9 (Fig. 51)
tapering, 3.6 times as long as its greatest diameter; proximal seta
very thin tapering pointed, distal seta cylindrical blunt striate,
proximal seta 0.3 of the length of tarsus and 1.7 times as long as the
length of distal seta. Cuticle of tarsus glabrous.

*Pygidium* (Fig. 52). *Tergum*. Posterior margin with three posterior lobes, one each posterior of insertion point of *st* and one broader in between. Relative lengths of setae, *a*_{1} not proved, *a*_{2}=10, *a*_{3}=23, *st*=15; *a*_{2} cylindrical blunt glabrous, *a*_{3} tapering pointed, striate distally, both setae curved inward and diverging; *st* somewhat clavate striate, directed posteriorly and curved inward. Distance *st-st* 2.2 times as long as *st*.

*Sternum*. Posterior margin with low lobe inside each seta *b*_{1}, straight in between. Relative lengths of setae (pygidial *a*_{2}=10): *b*_{1}=37, *b*_{2}=21, setae thin tapering, *b*_{1} striate, as long as interdistance, *b*_{2} almost as long as distance *b*_{1}-*b*_{2}.

Anal plate directed posteriorly, 1.2 times as long as wide, linguiform with convex lateral and posterior margins, thickened in most posterior part, two short appendages protruding from the middle of tergal side.

Figures 46–52.

*Decapauropus undulatus* sp. n., holotype, ad. 9(♂) **46** right antenna, sternal view **47** collum segment, median and left part, sternal view **48** tergite VI, posterior part **49** *T*_{3}**50** seta on trochanter of leg 9 **51** tarsus of leg 9 **52** pygidium, posteromedian and left part, sternal view. Scale: a: Figs 49–51; b: 47, 48; c: Fig. 52, d: Fig. 46.

Diagnosis.

A genus in Pauropodidae
with numerous and irregularly placed setae on tergites I-V, legs 1–9
5-segmented and interposed pairs 6-segmented, pygidial sternum with
setae *b*_{1} and *b*_{2}.

Etymology.

The genus name referring to the site of collecting at Doi Inthanon, formerly known as Doi Angka.

Genotype.

*Angkapauropus leptotrichos* sp. n.

urn:lsid:zoobank.org:act:90CF2C6D-2166-46A1-B905-74DE0B07E199

http://species-id.net/wiki/Angkapauropus_leptotrichos

Figs 53–60
Material.

Holotype ad. 9(♀), Thailand, Chiang Mai province, Doi Inthanon, Mae Chaem road, secondary dry forest, litter, alt. 1150 m, loc. CM-084. – Paratype, ad. 9(♀), ibidem, loc. CM-086. – 2 specimens.

Etymology.

From the Greek *leptos*, thin, and *trichos*, hair (referring to the shape of the setae on head, antennae and legs.

Description.

*Length.* 0.54(0.56) mm. *Head* (Fig. 53). Setae on tergal side short thin cylindrical striate. Relative lengths of setae (holotype only), 1^{st} row: *a*_{1}=10, *a*_{2}=13; 2^{nd} row: *a*_{1}=?, *a*_{2}=13, *a*_{3}=10; 3^{rd} row: *a*_{1}=7, *a*_{2}=13; 4^{th} row: *a*_{1}=13, *a*_{2}*=*17, *a*_{3}=20, *a*_{4}=13; lateral group setae not studied. Ratio *a*_{1}*/a*1*-a*1
in 1^{st} row 0.6, 2^{nd} row ?, 3^{rd} row 0.2, 4^{th} row 0.4. Temporal organs
large, in tergal view ovoid, length as long as their shortest
interdistance. Pore not ascertained. Head cuticle glabrous.

*Antennae* (Fig. 54). Segment 4 with five cylindrical striate setae; their relative lengths: *p*=10, *p’*=(8-)9, *p’’=r=*5, *p’’’*=3. Tergal seta *p* 0.6(-0.7) of the length of tergal branch *t*.
The latter fusiform with distal end cut obliquely, (2.4-)2.6 times as
long as its greatest diameter and (1.0-)1.2 times as long as the length
of sternal branch*s*, that branch 1.6 times as long as its greatest diameter; anterodistal corner distinctly truncate. Seta *q* cylindrical striate, almost as long as the length of *s*. Relative lengths of flagella (basal segments included) and basal segments: *F*_{1}=100, *bs*_{1}=9(-10), *F*_{2}=45(-52), *bs*_{2}=4(-5); *F*_{3}=87(-91), *bs*_{3}=5, curved. *F*_{1} 4.2(-4.7) times as long as *t*, *F*_{2} and *F*_{3} 1.8 and 4.4(-4.5) times as long as *s* respectively. Distal calyces conical, distal part of flagella axes not widened below calyces. Globulus *g* shortly pyriform, as long as wide, ≈10 bracts, capsule somewhat flattened; width of *g* as long as the greatest diameter of *t*. Antennae glabrous.

*Trunk* (Figs 55, 56, 60). Setae of collum segment (Fig. 55)
short simple cylindrical annulate, sublateral setae 1.2 times as long
as submedian setae; sternite process triangular; appendages cylindrical,
diameter of caps distinctly shorter than top of appendage. Process and
appendages glabrous.

Setae on tergites short cylindrical, numerous and irregularly inserted on I-V, ≈40 setae on I (Fig. 56),
≈15 on V, setae on VI in two transversal rows, 6 ones in anterior
row and 4 ones in posterior row. Submedian posterior setae on VI (Fig. 60) 0.1 of interdistance and as long as the length of pygidial setae *a*_{1}.

*Bothriotricha* (Fig. 57). Relative lengths: *T*_{1}=100, *T*_{2}=?, *T*_{3}=74(-80), *T*_{4}=91(-105), *T*_{5}=117(-124); axes simple straight, very thin and with faint pubescence only distally on *T*_{1}, *T*_{2} and *T*_{5}, *T*_{3} only (Fig. 57) stronger, with longish cylindrical end-swelling, 1/3 of the length of bothriotrix, distinct but sparse oblique pubescence.

*Legs* (Figs 58, 59). Setae on coxa (Fig. 58) and trochanter of legs 1–9 simple thin cylindrical annulate. Tarsus of leg 9 (Fig. 59)
subcylindrical, only somewhat tapering, 3.0 times as long as its
greatest diameter; setae very thin, cylindrical annulate, proximal
seta 0.1 of the length of tarsus and as long as distal seta. Cuticle of
tarsus glabrous.

*Pygidium* (Fig. 60). *Tergum*. Posterior margin with distinct posterior lobes outside setae *st, * in between straight. Relative lengths of setae: *a*_{1}=*st*=1, *a*_{2}=3, *a*_{3}=15. Setae very different, *a*_{1} short clavate, *a*_{2} short lanceolate, diverging, *a*_{3} long tapering, curved inwards, diverging, *st* short clavate, converging. Distance *a*_{1}*-a*1 (5.7-)6.7 times as long as the length of *a*_{1}, distance *a*_{1}*-a*2 (3.2-)3.7 times as long as distance *a*_{2}*-a*3; distance *st-st* 8(-9) times as long as *st* and 1.1(-1.5) times as long as distance *a*_{1}*-a*1.

*Sternum*. Posterior margin almost straight between insertion areas of setae *b*_{1}, no lobe below anal plate. Relative lengths of setae (pygidial *a*_{1}=1): *b*_{1}=9(-10), *b*_{2}=3, setae tapering pointed, glabrous, *b*_{1} 0.6(-0.9) of the length of interdistance, *b*_{2} 0.6 of the length of distance *b*_{1}-*b*_{2}.

Anal plate broadest anteriorly, distinctly longer than broad, lateral margins concave, each posterior corner with long thin subcylindrical appendage protruding backward and somewhat curved inward; between these appendages a shorter one protruding backward from tergal side, long appendages almost as long as plate, shorter one 0.7 of the length of plate.

Figures 53–60.

*Angkapauropus leptotrichos* gen. n., sp. n., holotype, ad. 9(♀) **53** head, median and right part, tergal view **54** right antenna, sternal view **55** collum segment, median and left part, sternal view **56** tergite I, right half **57** *T*_{3}**58** seta on coxa of leg 9 **59** tarsus of leg 9 **60**
posterior part of tergite VI and pygidium, posteromedian and left
posterior corner, tergal view. Scale: a: Fig. 56; b: 53, 55, 57–59;
c: 54. 60.

Genus

urn:lsid:zoobank.org:act:786F9B4C-205F-44C5-B98F-99128031C94C

http://species-id.net/wiki/Borneopauropus_platylopas

Figs 61–71
Material.

Holotype ad. 9(♂), Thailand, Chiang Mai province, Doi Inthanon, Mae Chaem road, secondary dry forest, litter, alt. 1150 m, 1992.vi.30, loc. CM-080. – Paratype, ad. 9(♀), ibidem, loc. CM-194. – 2 specimens.

Etymology.

From the Greek *platys*, broad, and *lopas*, plate (referring to the unusually broad anal plate).

Diagnosis.

Up to now four species have been described in
the genus, one from Sulawesi in Indonesia, two from Sabah and one
from Tasmania. *Borneopauropus platylopas* has most characters in common with the Indonesian species, *Borneopauropus curtipes* Scheller (*Borneopauropus penanorum* Scheller (*T*_{3}, clavate part moderately widened with long distal flagellum in *Borneopauropus platylopas*, not strongly widened and with shorter flagellum in *Borneopauropus curtipes* and *Borneopauropus penanorum*, and by the shape of the pygidial setae *a*_{3}, long thin tapering, not bladder-shaped, and the shape of the anal plate, broad triangular, not longish linguiform.

*Description. Length.* (0.51-)0.58 mm. *Head* (Fig. 61). Tergal and lateral sides with 26 setae arranged as in Fig. 61;
transversal rows indistinct laterally. Setae longest in posteromedian
part; all but lateral setae clavate, shortly pubescent, lateral group
setae cylindrical annulate. Temporal organs small with at least two
short uplifted extensions. Head cuticle glabrous.

*Antennae* (Fig. 62). Segment 4 with 4 cylindrical annulate setae; relative lengths: *p*=10, *p´*=11(-12), *p´´*=4(-6), *r*=4. Tergal seta *p* 0.6 of the length of tergal branch *t*. The latter subcylindrical, (2.6-)2.9 times as long as wide, 0.9 of the length of sternal branch *s*. That branch thickest in distal third and with anterodistal corner somewhat more truncated than posterodistal corner; *s* (2.0-)2.1 times as long as greatest diameter, its seta *q* cylindrical striate, 0.5 of the length of *s*. Relative lengths of flagella (base segments included) and base segments: *F*_{1}=100, *bs*_{1}=8; *F*_{2}=(65-)66, *bs*_{2}=6; *F*_{3}=72(-74), *bs*_{3}=(7-)8. *F*_{1} 3.1 times as long as *t, F*2 and *F*_{3} (1.9-)2.2 and (2.0-)2.3 times as long as *s* respectively. Distal calyces helmet-shaped, glabrous. Globulus *g*, (1.6-)1.8 times as long as greatest diameter, the latter (0.9-) as long as greatest diameter of *t*. Antenna glabrous.

*Trunk* (Figs 63-65). Setae of collum segment (Fig. 63)
simple cylindrical blunt annulate; sublateral seta 1.2 times as long as
submedian seta. Sternite process blunt-ended; appendages barrel-shaped
with low caps; process and appendages glabrous.

Tergites (Figs 64, 65), I, V, VI entire, II, III, IV transversely and weakly 2-parted. Number of setae on tergites (holotype only, if two groups of values anterior and posterior groups of setae respectively): I 30, II 19+20, III 28+21, IV 25+19, V 20+13, VI 4+2. Setae bladder-shaped with short oblique-erect pubescence, setae lengthening posteriorly, those on VI (Fig. 65) about twice longer than those on I. Cuticle of tergites glabrous.

*Bothriotricha* (Figs 66, 67). Relative lengths: *T*_{1}=100, *T*_{2}=108(-110), *T*_{3}=(122-)146, *T*_{4}=(89-)113, *T*_{5}=(140-)144. Axes simple, most proximally glabrous; proximal half of *T*_{3} (Fig. 66) strongly clavate with short pubescence arranged in dense whorls, distal half very thin; other bothriotricha (Fig. 67) with thin and curved axes; pubescence short oblique on proximal parts, erect distally.

*Genital papillae* (Fig. 68). Short, as long as wide, seta 0.5 of the length of papilla.

*Legs* (Figs 69, 70). All legs 5-segmented. Setae on coxa (Fig. 69)
and trochanter of legs 1–9 simple, cylindrical annulate blunt,
rudimentary secondary branch only on coxal setae of leg 2 in male.
Tarsus of leg 9 (Fig. 70)
short, almost cylindrical, (2.9-)3.4 times as long as greatest
diameter; setae short cylindrical annulate, proximal one 0.1 of the
length of tarsus and (0.5-)0.6 of the length of distal seta. Cuticle of
tarsus glabrous.

*Pygidium* (Fig. 71)*. Tergum*. Hind margin rounded. Relative lengths of setae: *a*_{1}=10, *a*_{2}=(13-)14, *a*_{3}=(22-)25, *st* rudimentary. *a*_{1} bladder-shaped, faintly pubescent, *a*_{2} longish clavate, distinctly pubescent, somewhat curved inward, *a*_{3} thin tapering, faintly striate, curved inward. Distance *a*_{1}–*a*_{1} 1.5 times as long as *a*_{1}, distance *a*_{1}–*a*_{2} twice longer than distance *a*_{2}–*a*_{3}; distance *st*–*st* ≈25 times as long as *st* and 0.6 of distance *a*_{1}–*a*_{1}.

*Sternum*. Posterior margin between *b*_{1}
rounded but with large lobe below anal plate; lobe with median
indentation and rounded posterolateral corners. Relative lengths of
setae (pygidial *a*_{1}=10): *b*_{1}=(40-)43, *b*_{2}=13(-16), *b*_{3}=7. *b*_{1} thin tapering, blunt, striate distally, *b*_{2} and *b*_{3} cylindrical. *b*_{1} (1.3-)1.4 times as long as distance *b*_{1}-*b*_{1}*, b*2 0.7(-0.9) of distance *b*_{1}-*b*_{2} and *b*_{3} 0.3(-0.4) of distance *b*_{3}-*b*_{3}.

Anal plate (Fig. 71) narrowest anteriorly, triangular, posterolateral corner turned anteriorly and posterior margin with small median process, two cylindrical blunt faintly pubescent appendages protruding backward from sternal side just nearby the posteromedian process, length of appendages somewhat longer than plate.

Figures 61–71.

*Borneopauropus platylopas*, sp. n., holotype, ad. 9(♂) **61** head, median and right part, tergal view **62** right antenna, tergal view **63** collum segment, median and left part, sternal view **64** tergites I-II, right half **65** tergite VI, right half **66** *T*_{3}**67** *T*_{2}**68** genital papillae and seta on coxa of leg 2, anterior view **69** seta on coxa of leg 9 **70** tarsus of leg 9 **71** pygidium, posterior part, tergal view. Scale: a: Figs 64, 65; b: Figs 66–70; c: Fig. 61; d: 62, 63; e: 71.

Genus

Material.

Thailand, Chiang Mai province, Doi Inthanon, Mae Chaem road, secondary dry forest, litter, alt. 1150 m, 1991.vi.30, 1 ad. 9(♀), ibidem, litter, 1991.vi.30, loc. CM-074, ibidem, litter, 1 ad. 9(♂), 1991.vi.30, loc. CM-080, ibidem, litter, 1 juv. 6, 30.vi.1991, CM-089; below the top, alt 2400, 1 ad. 9(♀), 1991.vii.4, loc. CM-202. – 4 specimens.

General distribution.

Known from Doi Inthanon only (

Material.

Thailand, Chiang Mai province, Doi Inthanon, secondary dry forest, litter, alt. 1150 m, 1 ad. 9(♀), 1991.vi.30, loc. CM-086. – 1 specimen.

General distribution.

Known from Doi Inthanon only (

urn:lsid:zoobank.org:act:CC141A91-BC0A-4E8B-B550-BDBE60F9D981

http://species-id.net/wiki/Samarangopus_choanephorus

Figs 72–83
Material.

Holotype ad. 9(♂), Thailand, Chiang Mai province, Doi Inthanon, below the top, primary rain forest, soil, alt. 2400 m, 1991.vii.4, loc. CM-195. – 1 specimen.

Etymology.

From the Greek *choane*, funnel, and *phero*, bear, carry (referring to the funnel-shaped organs on the tergites).

Diagnosis.

*Samarangopus choanephorus* sp. n. may be grouped together with *Samarangopus umbonifer* Scheller from Thailand (*Samarangopus condylus* Scheller from the Philippines (*Samarangopus umbonifer* are the shape of the antennal globulus *g*, stalk much longer than globulus in *Borneopauropus choanephorus*, shorter in *Borneopauropus umbonifer*,
the large funnel-shaped organs of the tergites, with central rod,
not without, the protuberances of the posterior margin of the tergite
VI, with evenly convex lateral margins, not with a small knob on each
side, the shape of the setae on coxa and trochanter of leg 9, simple,
not furcate, and the length of the pygidial setae *b*_{3}, almost as long as the *b*_{2}, not distinctly shorter.

Description.

*Length.* 0.74 mm. *Antennae* (Fig. 72). Chaetotaxy of segments 1–4: 2/2/2/2; no *g´*. Setae thin striate, on segment 4 only two proved, their relative lengths *p*=10, *p´*or *p´´*=6. Tergal branch *t* almost cylindrical, 3.1 times as wide as greatest diameter and as long as sternal branch *s*. That branch twice longer than greatest diameter, anterodistal corner distinctly truncate. Seta *q* as setae of 4^{th} segment, 0.5 of the length of *s.* Relative lengths of flagella (base segments included) and base segments: *F*_{1}=100, *bs*_{1}=18, *F2*=?, *bs*2~10, *F3*=85, *bs*_{3}=17. *F*_{1} 5.6 times as long as *t*, *F3* 3.3 times as long as *s*. Calyces conical; flagella axes inconsiderably widened below calyx. Globulus *g* 1.6 times as long as greatest width; 10 thin bracts, capsule spherical, diameter of *g* 0.8 of the greatest diameter of *t.* Antenna glabrous.

*Trunk* (Figs 73–77). Setae of collum segment (Fig. 73)
similar thin furcate, main branch pointed striate, secondary branch
rudimentary pointed; sublateral setae 1.3 times as long as submedian
setae. Sternite process broad triangular pointed anteriorly. Appendages
wide cylindrical, caps flat. All parts of collum segment glabrous.

Tergites with four main types of protuberances (Figs 74–77): 1. stalked campanulate marginal protuberances (Figs 74–77), short on anterior margin of tergite I, larger on lateral margins of tergites; 2. fungiform organs (Figs 74–77)
with hat in the shape of an upside down transparent funnel with a
central rod, foot subcylindrical, these distributed over whole the
surface of tergites; 3. small subcylindrical organs with upside down
funnel at top (Fig. 77);
4. many small conical structures in between the different
protuberances. Number of marginal protuberances: (holotype only): I –
36, II, 1 small – *T*_{1} – 11; III, 8 – *T*_{2} – 8; IV, 8 – *T*_{3} – 6; V, 9 – *T*_{4} – 4, VI, 7 – *T*_{5} – 1.

*Bothriotricha.* All but *T*_{3} with very thin axes, curved distally. *T*_{3} (Fig. 78)
with thicker axes distally forming a subcylindrical, almost glabrous,
swelling, length ¼ of the length of bothriotrix. Relative lengths of
bothriotricha (holotype only): *T*_{1}=100, *T*_{2}=95, *T*_{3}=42, *T*_{4}≈90, *T*_{5}=89.

*Genital papillae* (Fig. 79).
Conical, glabrous, 2.2 times as long as greatest diameter, setae
long, 0.8 of the length of papillae, inserted below the middle of
papillae.

*Legs* (Figs 80, 81).All legs 5-segmented. Seta on coxa and trochanter (Fig. 80) of legs 1–9thin simple striate. Tarsi tapering, those of leg 9 (Fig. 81)
3.7 times as long as its greatest diameter; two tergal setae, both
pointed glabrous; length of proximal one 0.4 of the length of tarsus and
3 times longer than distal seta. Cuticle of tarsus glabrous. No
proximal seta on tarsi of leg 1. All legs with large main claw and small
setose anterior secondary claw; in leg 9 the former reaching 0.5 of the
length of tarsus. Appendage on femur of leg 1 not studied.

*Pygidium* (Figs 82, 83). *Tergum.* Posterior margin undulate with larger pentagonal lobe between *st.* Setae *a*_{1} cylindrical, curved inward, converging, *a*_{2} short clavate straight converging, faintly pubescent distally, *a3* long thin tapering pointed diverging, *st* straight lanceolate, faintly pubescent. Relative lengths of setae: *a*_{1}=10, *a*_{2}=6(-7), *a*_{3}=(20-)21, *st*=(15-)16. Distance *a*_{1}-*a*_{1} 1.3 times as long as *a*_{1}, distance *a*_{1}-*a*_{2} as long as distance *a*_{2}*–a3*; distance *st*-*st* almost as long as *st* and 1.1 times as long as distance *a*_{1}-*a*_{1}. Cuticle glabrous.

*Sternum* (Fig 83). Posterior margin rounded. Setae long tapering pointed, *b*_{2} straight diverging, *b*_{3} somewhat curved inward. Relative lengths of setae (pygidial *a*_{1}=10): *b*_{1}=(45-)47, *b2*=(26-)29, *b3*=24(-25). *b*_{1} 1.1 times as long as interdistance, *b*_{2} 1.1 times as long as distance *b*_{1}*-b*2, *b*_{3} 0.6 of interdistance.

Anal plate (Fig. 82) 1.4 times as long as broad, lateral margins convex, distal part of plate cleft by a V-shaped incision into two somewhat tapering cylindrical branches; four appendages: each branch provided with one outer and one inner tooth and in between a straight bladder-shaped appendage, that broadest in distal half, length of appendage 0.4 of the length of plate; two short thin cylindrical appendages protruding outward-backward from lateral margins. Plate glabrous, appendages with faint pubescence.

Figures 72–83.

*Samarangopus choanephorus* sp. n., holotype ad. 9(♂) **72** left antenna, sternal view **73** collum segment, median and left part **74** tergite I, anterolateral margin with campanulate protuberances and fungiform organs **75** tergite I, posterolateral corner **76** tergite IV, right lateral margin around insertion pit of *T*_{3}**77** tergite VI, right posterolateral part **78** *T*_{3}**79** genital papillae, anterior view **80** seta on trochanter of leg 9 **81** tarsus of leg 9 **82** pygidial tergum, posterior part, tergal view **83** pygidial sternum, sternal view. Scale: a: Fig. 79; b: Figs 74–76; c: Figs 72, 73, 77, 78, 80, 81; d: Figs 82, 83.

Material.

Thailand, Chiang Mai province, Doi Inthanon, below the top, litter, alt. 2400 m, 1 ad. 9(♀), 1991.vii.4, loc. CM-220.

General distribution.

Known from Doi Inthanon only (** **

urn:lsid:zoobank.org:act:980E30A4-B446-46AA-8887-A487ACC4B385

http://species-id.net/wiki/Sphaeropauropus_angulatus

Figs 84–94
Material.

Holotype ad. 9(♂), Thailand, Chiang Mai province, Doi Inthanon, near Mae Chaem crossing, litter, alt. 1700 m, 1991.vii.12, loc. CM-162. – Paratype ad. 9(♂), below the top, primary rain forest, litter, alt. 2400 m, 1991.vii. 4, loc. CM-206. – Non type material, Mae Chaem road, secondary dry forest, litter, alt. 1150 m, 1juv. 3, 1991.vi.30, loc. CM-064, ibidem, litter, 1 subad. 8(♀), 1991.vii.4, loc. CM-096, ibidem, below the top, primary rain forest, litter, alt. 2400 m, 1 juv. stad.?, 1991.vii.4, loc.CM-194, ibidem, below the top, primary rain forest, soil, 1 subad. 8(♀), 1991.vii.4, loc. CM-195, ibidem, litter, 1 ad. (♀), 1991.vii.4, loc. 202, ibidem, soil, 1 stad.?, 1991.vii.4, loc. CM-207, ibidem, litter, 1 juv. 5, 1 juv. 3, 1991.vii.4, loc. CM-212, ibidem, litter, 1 ad. 9(♀), 1991.vii.4, loc. CM-219, ibidem, litter, 2 juv. stad?, 1991.vii.4, loc. CM-222. – 13 specimens.

Etymology.

From the Latin *angulatus*, with angles (referring to the shape of the lateral margins of the anal plate).

Diagnosis.

*Sphaeropauropus angulatus*
is well defined from its congeners by the shape of the anal plate with
distinct lateral corners and by the cuticular pattern of the tergites.
At present its relationships can not be traced.

Description.

*Antennae* (Fig. 84). Chaetotaxy of segments 1–4: 2/2/2/3. Setae of first three segments and seta *p*’’ of 4^{th} segment thin cylindrical, faintly striate, *p* and *p*’ of 4^{th} segment distinctly tapering, pointed glabrous, relative length of setae there: *p*=10, *p*’=*p*’’=10, *r=*5. *g´* of 3^{rd} segment not proved. Tergal seta *p* 0.8 of the length of tergal branch *t.* The latter cylindrical, 4.0 times as long as the greatest diameter and 1.1 times as long as sternal branch *s.* That branch subcylindrical, 2.8 times as long as the greatest diameter. Anteriodistal truncation small, seta *q* short thin pointed, 0.2 of the length of *s.* Relative lengths of flagella (base segments included) and base segments: *F*_{1}=100, *bs*_{1}=13, *F*_{2}=51, *bs*_{2}=7, *F*_{3}=89, *bs*_{3}=15. *F*_{1} 2.8 times as long as *t, F*2 and *F*_{3} 1.5 and 2.7 times as long as *s* respectively. Globulus *g* straight, stalk thin, *g*
2.2 times as long as its greatest diameter; ≈10 bracts, capsule almost
spherical. Calyces small conical, flagella axis not widened below
calyx. Cuticle of antennae glabrous.

*Trunk* (Figs 85–89). Submedian setae of collum segment (Fig. 85)
not proved, sublateral setae thin simple striate; sternite process
broad low; appendages short cylindrical with thin caps. Process and
appendages glabrous.

Tergites with tuft-like setae with clavate stalk (Figs 86–88). Cuticle between setae (Fig. 89) with very small sessile organs in the centre of round glabrous spots surrounded by dense short pubescence arranged in a checkered pattern. Pubescence longest on lateral parts of tergites.

*Bothriotricha*. All but *T*_{3} with very thin glabrous, very shortly pubescent axes. *T*_{3} (Fig. 90)
with thicker axes and distal swelling, length of the latter 1/3 of the
length of bothriotrix. Relative lengths of bothriotricha: *T*_{1}=100, *T*_{2}=144, *T*_{3}=62, *T*_{4}=139, *T*_{5}=146.

*Genital papillae* (Fig. 91).
Longish, 3.4(-3.6) times as long greatest diameter, proximal half
subcylindrical, distal half tapering, seta 0.3 of the length of
papilla. Cuticle glabrous. Coxal seta of leg 2 as on leg 1.

*Legs* (Figs 92, 93). All legs 5-segmented. Setae on coxa and trochanter of legs 9 (Fig. 92)
furcate, main branch very thin, striate; secondary branch rudimentary
pointed glabrous. These setae on legs 1–8 similar. Tarsus of leg 9
slender bow-shaped, (4.9-)5.0 times as long as greatest diameter, two
tergal setae, both tapering pointed glabrous. Proximal seta 0.2 of the
length of tarsus and (4.0-)4.2 times as long as distal seta. No proximal
seta on tarsus of leg 1. Appendage of anterior side of femur of leg 1
not studied. Legs glabrous. All legs with almost straight main claw and
small setose anterior secondary claw, length of main claw 0.3 of the
length of tarsus.

*Pygidium* (Fig. 94)*. Tergum*.
Posterior margin straight. Setae subsimilar to each other, cylindrical
straight glabrous, directed posteriorly. Relative lengths of setae: *a*_{1}=10, *a*_{2}=6(-7), *a*_{3}=14(-16), *st*=(11-)12. Distance *a*_{1}-*a*_{1} 2.5 times as long as *a*_{1}, distance *a*_{1}-*a*_{2} almost as long as distance *a*_{2}*–a3*; distance *st*-*st* 1.6 times as long as *st* and 0.5 of distance *a*_{1}-*a*_{1}. Cuticle glabrous.

*Sternum.* Posterior margin straight. Setae tapering pointed, *b*_{3} converging. Relative lengths of setae (pygidial *a*_{1}=10): *b*_{1} lost, *b2*=13, *b3*=16. *b*_{2} as long as distance *b*_{1}*-b*2, *b*_{3} 0.3 of interdistance.

Anal plate pentagonal with distinct lateral corners and two somewhat diverging cylindrical branches protruding backwards from posterior part of sternal side; each branch with two distal appendages: one inner short pointed, the other long clavate; the latter 0.5 of the length of plate; clavate appendages faintly pubescent, other parts of plate glabrous.

Figures 84–94.

*Sphaeropauropus angulatus* sp. n., holotype ad. 9(♂) **84** left antenna, sternal view; 85 collum segment, median and left part **86** tergite I, anterior margin, sternal view **87** tergite I, anterolateral margin, tergal view **88** tergite VI, posteromedian margin, sternal view **89** tergite I, inner part, tergal view **90** *T*_{3}**91** genital papillae, anterior view **92** seta on coxa of leg 9 **93** tarsus of leg 9 **94** pygidium posteromedian and left part, sternal view, only insertion points of *b*_{1} shown. Scale: a: Fig. 91; b: Figs 86–88, 92; c: Fig. 93; d: Fig. 94; e: Figs 84, 85, 89, 90.

Pauropods have been reported from Thailand twice only, by *Allopauropus* and *Decapauropus* (Pauropodidae), and by the present author (*Decapauropus* in Pauropodidae, *Samarangopus* in Eurypauropodidae, and *Sphaeropauropus* in Sphaeropauropodidae. Now four species described earlier and ten new species are reported: six ones in *Decapauropus* and one in the new genus *Angkapauropus* in Pauropodidae, one in *Borneopauropus* in Brachypauropodidae, a family new to Thailand, one species in *Samarangopus* in Eurypauropodidae, and one in *Sphaeropauropus*
in Sphaeropauropodidae. Three of the new species described above seem
to be related to species occurring in south east Asia (Indonesia, the
Philippines, Sabah) but no less than five species have specific
characters or character combinations which, as far as now known,
indicate that they belong to a fauna with restricted geographic
distribution.

Together 21 species are known from Doi Inthanon representing six genera in four families. Only one of the species, *Decapauropus mortensenii*
Hansen, has been collected outside, in Thailand on the Island Koh
Chang and also in a large area from Egypt in the west to New Caledonia
in the east.

From the whole Thailand 28 species in four families and six genera are now known. Because only one of these species has a wide range the degree of endemism seems to be unusually large.

References

Hansen
HJ (1902) On the genera and species of the order Pauropoda.
Videnskabelige Meddelelser fra den naturhistoriske Forening i
Kjøbenhavn, for aaret 1901: 323–424, pls I-VI.

Scheller U (1990) Pauropoda and Symphyla
(Myriapoda) collected on St. John, U.S. Virgin Islands. Caribbean
Journal of Science 25 (3–4):164-195.

Scheller U (1995) Pauropoda (Pauropodidae Eurypauropodidae) from north-western Thailand. Tropical Zoology 8 (1):7-41.

Scheller U (2009) Records of Pauropoda
(Pauropodidae, Brachypauropodidae, Eurypauropodidae) from Indonesia
and the Philippines with description of a new genus and 26 new species.
International Journal of Myriapodology 2:69-148. doi: 10.1163/187525409X12577705044548

Scheller U, Brinck P, Enckell PH (1994) First record of Pauropoda (Myriapoda) from Borneo. Stobaeana 1:1-14.